LINCOLN, Neb. (AP) — New Nebraska football coach Mike Riley has hired four assistant coaches, leaving him with one spot to fill on his staff.

The assistants named Wednesday are running backs coach Reggie Davis, defensive line coach Hank Hughes, offensive coordinator Danny Langsdorf and secondary coach Charlton Warren.

Langsdorf was the New York Giants quarterbacks coach this season after working nine years as Riley’s offensive coordinator and quarterbacks coach at Oregon State.

Davis was an assistant with the San Francisco 49ers. Hughes was co-defensive coordinator and linebackers coach at the University of Cincinnati. Warren, hired by former Nebraska coach Bo Pelini, is being retained.

Riley has already hired defensive coordinator Mark Banker, linebackers coach Trent Bray, offensive line coach Mike Cavanaugh and special teams coordinator Bruce Read.